Part III
Back to our Maximum Likelihood estimate of r. The general
formula is:
L = C + a1 log m1 + a2 log m2 +...+ ai log mi
We take the first derivative of L with respect to r
and set this equal to zero to maximize r.

| |
Numbers |
| Classes |
Observed |
Expected |
| PpTt |
a1 |
m1
= (1- r)/2 |
| pptt |
a4 |
m4
= (1- r)/2 |
| Pptt |
a2 |
m2
= r/2 |
| pptt |
a3 |
m3
= r/2 |
We have shown that m = 1/2(1-r) where we multiply 1/2(1-r)
for a single event by n to get the total number expected
for n progeny.

Because we substituted m = (1 - r)/2, etc.
n/2 is a constant so when q =
a
log (1-r)
dq
= - a because
d log (1-r) =
- 1
dr 1-r because
d lodr(1-r) = -
1-r
