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Jerzy A. Lis
Department of Biosystematics,
University of Opole, Poland

     I am a Professor in the Department of Biosystematics at the University of Opole. My research concentrates on the systematics of the Pentatomoidea, especially the morphology, taxonomy, classification, and phylogeny of the Cydnidae and related families (the so called “cydnoid-complex”). For a long time, much of my research has focused on the taxonomy of the family Cydnidae; During my revisional studies I have described and named about 200 new taxa of the Cydnidae (more than 180 new species and about 20 new genera). I have revised also the Cydnidae in the Oriental region and prepared a catalogue of the Old World Cydnidae. At present I concentrate on comparative studies of different structures in Cydnidae and related families trying to evaluate their significance for the taxonomy, phylogeny and classification of the Cydnidae and the whole “cydnoid-complex”, as well (including projects with Dominique Pluot-Sigwalt of Paris, and Carl W. Schaefer of Connecticut). All these studies aim in the reclassification of the family.

     I am also interested in the taxonomy and morphology of Dinidoridae, and I study the Heteroptera fauna of Poland, as well.
 

     Moreover, I am a leader of the team working on the Heteroptera at the University (the largest team in Poland): Anna Kocorek (Dinidoridae), Barbara Lis (Cimicomorpha, esp. Tingoidea; faunistics of Poland), Anna Hohol-Kilinkiewicz (Tingidae, Anthocoridae, Cydnidae), Joanna Czaja (Scutelleridae, Cydnidae), Grzegorz Hebda (faunistics of Poland).

Contact Information:
Department of Biosystematics
Division of Invertebrate Zoology
University of Opole
Oleska 22, 45-052 Opole
POLAND

cydnus@uni.opole.pl

Current Projects:

1) Morphological diversity of the spermatheca among Cydnidae (Heteroptera) and its bearing on the classification of the family (with Dominque Pluot-Sigwalt).

2) Comparative studies on the coxal combs in the Cydnidae and related families (with Carl W. Schaefer).

3) Comparative studies on the tarsus and pretarsal structures in the Cydnidae and related families.

4) External morphology of the abdominal lateral sclerites of the Cydnidae (with Anna Hohol-Kilinkiewicz of my Department).

5) Comparative studies of the male and female genitalia in the Cydnidae.

6) A revision of the genus Geopeltus from New Caledonia, with new data on distribution and biology (with Geoff Monteith).

7) New species and new genera of Cydnidae from different regions.

 

Publications:

Lis, J. A. 1986. Carpocoris melanocerus (Mulsant et Rey, 1852) (Heteroptera, Pentatomidae) – a species of shield-bugs new to Poland. Polskie Pismo Entomologiczne 56: 221-222. [in Polish, with English summary]

Lis, J. A. 1987. Anomalies of the colouration and sculpture of the body of Dolycoris baccarum (L.) (Heteroptera, Pentatomidae). Polskie Pismo Entomologiczne 56[1986]: 917-918.

Lis, J. A. 1987. Bugs of the superfamily Pentatomoidea Reut. (Hemiptera: Heteroptera) of the Silesian Upland. Fragmenta faunistica 30: 359-372. [in Polish, with English summary] 

Lis, J. A. 1989. Shield-bugs of Poland (Heteroptera, Pentatomoidea) - a faunistic review. I. Plataspidae, Thyreocoridae, Cydnidae, Scutelleridae and Acanthosomidae. Polskie Pismo Entomologiczne 59: 27-83. 

Lis, J. A. 1989. New records of African Phyllocephalinae (Heteroptera, Pentatomidae). Polskie Pismo Entomologiczne 59: 393-395. 

Lis, J. A. 1990. Shield-bugs of Poland (Heteroptera, Pentatomoidea) - a faunistic review. Pentatomidae. Annals of the Upper Silesian Museum, Entomology 1: 5-102. 

Lis, J. A. 1990. New genera, new species, new records and checklist of the Old World Dinidoridae (Heteroptera, Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 1: 103-147. 

Lis, J. A. 1990. Studies on the Oriental Cydnidae. I. A revision of the genus Scoparipes Signoret, 1879 (Heteroptera). Annals of the Upper Silesian Museum, Entomology 1: 159-192. 

Lis, J. A. 1990. Studies on the Oriental Cydnidae.II. (Heteroptera). Three new genera and species. Bonner zoologische Beiträge 41(3-4): 223-229. 

Lis, J. A. 1991. Geotomus barbarae sp. n. from Madagascar and new records of some other Afrotropical burrowing bugs (Heteroptera, Cydnidae). Polskie Pismo Entomologiczne 60(3-4)[1990]: 173-176. 

Lis, J. A. 1991. On the last instar larvae of some Dinidoridae species (Heteroptera, Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 2: 83-92. 

Lis, J. A. 1991. Studies on Oriental Cydnidae. III. On the genus Peltoxys Signoret, 1880 (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 2(1): 19-25. 

Lis, J. A. 1991. Studies on Oriental Cydnidae. IV. New species, new synonyms and new records (Heteroptera). Annals of the Upper Silesian Museum, Entomology 2: 165-190. 

Lis, J. A. 1991. Studies on Oriental Cydnidae. V. Three new species of the genus Macroscytus Fieber (Heteroptera). Annals of the Upper Silesian Museum, Entomology 2: 209-215. 

Lis, J. A. 1991. Schiodtella secunda n. sp., a new species of burrowing bugs from Palaearctic China (Heteroptera: Cydnidae: Scaptocorinae). International Journal of Invertebrate Taxonomy - Genus 2(1): 27-31. 

Lis, J. A. 1991. Scoparipes Signoret, 1879 (Insecta, Heteroptera): Proposed confirmation of Cydnus latipes Westwood, 1837 as the type species. Bulletin of Zoological Nomenclature 48(3): 215-216. 

Lis, J. A. 1991. Studies on Oriental Cydnidae. VI. A contribution to the fauna of Sulawesi (Celebes) (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 2(4): 299-311. 

Lis, J. A. 1991. Studies on Oriental Cydnidae. VII. On Stibaropus Dallas and Pseudostibaropus n. gen. (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 2(4): 313-325. 

Lis, J. A. 1992. New records of Oriental and Australian Dinidoridae with descriptions of two new species (Heteroptera: Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 3: 35-43. 

Lis, J. A. 1992. On Garsauria aradoides Walker and G. laosana Lis (Heteroptera: Cydnidae). Annals of the Upper Silesian Museum, Entomology 3: 147-150. 

Lis, J. A. 1992. Studies on Oriental Cydnidae. IX. On the genus Adrisa Amyot and Serville, 1843 (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 3(1): 13-27. 

Lis, J. A. 1993. Studies on Cydnidae of the Australian Region. I. A review of species from Solomon Islands (Heteroptera: Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 4: 13-24. 

Lis, J. A. 1993. Studies on Cydnidae of the Australian Region. II. A review of New Guinean species of the genus Chilocoris Mayr (Heteroptera: Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 4: 25-40. 

Lis, J. A. 1993. Studies on Cydnidae of the Australian Region. III. Two new species of Macroscytus Fieb. from New Guinea (Heteroptera: Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 4: 41-46. 

Lis, J. A. 1993. A review of the tribe Amaurocorini E.Wagner with remarks on its systematic position within the family (Heteroptera: Cydnidae). Annals of the Upper Silesian Museum, Entomology 4: 59-78. 

Lis, J. A. 1993. On Byrsinus Fieber and Microporus Uhler, two allied cydnid genera (Heteroptera: Cydnidae). Annals of the Upper Silesian Museum, Entomology 4: 79-98. 

Lis, J. A. 1993. Microscytus, a new African genus of burrower bugs (Heteroptera: Cydnidae). Acta Entomologica Silesiana 1(1): 1-4. 

Lis, J. A. and W. Żyła. 1993. New record of Graphosoma lineatum (Linnaeus) (Heteroptera: Pentatomidae) in Upper Silesia. Acta Entomologica Silesiana 1(1): 17. [in Polish] 

Lis, J. A. 1993. Studies on Oriental Cydnidae. VIII. On Aethus indicus (Westwood), A. philippinensis Dallas and A. pseudindicus n. sp. (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 4(2): 103-111. 

Lis, J. A. 1993. Studies on Cydnidae of the Australian Region. IV. Two new species of Peltoscytus Lis from the Pacific islands (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 4(4): 303-308. 

Lis, J. A. 1994. A description of Parachilocoris minutus japonicus nov. subspec., and some changes in the nomenclature of East Palearctic burrower bugs (Heteroptera: Cydnidae). Entomologische Berichten 54(4): 113-116. 

Lis, J. A. 1994. A revision of Oriental burrower bugs (Heteroptera: Cydnidae). Upper Silesian Museum, Bytom: 349 pp. 

Lis, J. A. 1994. Studies on Cydnidae (Heteroptera) of the Australian Region. VI. Material collected by I. Lansbury in Papua New Guinea in 1990 and 1992. European Journal of Entomology 91: 321-325. 

Lis, J. A. 1994. Coleocydnus ernsti gen. et. sp.n., a coleopteroid representative of burrower bugs from tropical Africa, and new records of three recently described African species of the family (Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 5(3): 187-192. 

Lis, J. A. 1994. Studies on Cydnidae of the Australian Region. VII. On the genus Aethus Dallas, 1851 (Heteroptera: Pentatomoidea). International Journal of Invertebrate Taxonomy - Genus 5(4): 357-362. 

Lis, J. A. 1994. Studies on Oriental Cydnidae. X. Notes on the distribution of Cydnopeltus horvathii Signoret, and new records for two other species of the family (Heteroptera: Pentatomoidea). Annals of the Upper Silesian Museum, Entomology 5: 173-175. 

Lis, J. A. and J. Péricart. 1995. Scholtz ou Scholz: Note sur l'orthographe d'originelle du nom d'un Hétéroptériste silésien du XIXe sičcle. Bulletin de la Societe entomologique de France 99(5)[1994]: 482.

Lis, J. A. 1995. Studies on Cydnidae of the Australian Region. V. The genus Chilocoris Mayr in Australia, Tasmania and Moluccas (Heteroptera). Bonner zoologische Beiträge 45(3-4)[1994]: 225-230. 

Lis, J. A. 1995. A synonymic list of burrower bugs of the Australian Region (Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 6(2): 137-149. 

Lis, J. A. 1995. Studies on Oriental Cydnidae (Heteroptera). XI. A new species of Paraethus Lis from Java, with remarks on the systematic position of the genus. Zoologische Mededelingen 69(4): 53-56. 

Lis, J. A. 1995. The genus Macroscytus Fieber (Heteroptera: Cydnidae) in the East Palaearctic subregion. Entomologische Berichten 55(10): 163-165. 

Lis, J. A. 1996. Does the New World genus Dinidor Latreille (Heteroptera: Dinidoridae) occur in Africa? Acta Entomologica Silesiana 3(1-2)[1995]: 27-29. 

Lis, J. A. 1996. A review of burrower bugs of the Australian Region, with a discussion on the distribution of the genera (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 7(2): 177-238. 

Lis, J. A. 1996. Studies on Cydnidae of the Australian Region. VIII.  Cydnus (Orientocydnus) pericarti n. sp. from Queensland (Heteroptera). Bulletin de la Sociéte entomologique de France 101(1): 27-30. 

Lis, J. A. 1996. Studies on Cydnidae of the Australian Region. IX. Chilocoris cavitus n. sp. and Ch. solomonensis n. sp. from Solomon Islands (Hemiptera: Heteroptera). Annals of the Upper Silesian Museum, Entomology 6-7: 89-93. 

Lis, J. A. 1996. Lectotype designations for some Palaearctic Cydnidae (Hemiptera: Heteroptera). Annals of the Upper Silesian Museum, Entomology 6-7: 105-106. 

Lis, J. A. 1996. Paraethus sigwalti n. sp. from Senegal, and notes on Aethus (Stilbocydnus) laevis Wagner, 1951 (Hemiptera: Heteroptera: Cydnidae). Annals of the Upper Silesian Museum, Entomology 6-7: 247-248. 

Lis, J. A. and A. Kocorek. 1996. Thalma secunda n. sp. from New Guinea, and notes on other species of the family (Hemiptera: Heteroptera: Dinidoridae). Annals of the Upper Silesian Museum, Entomology 6-7: 249-252. 

Lis, J. A. and S. Skórka. 1996. On the systematic position of Calliphara bipunctata Lehmann, 1920 (Hemiptera: Heteroptera: Scutelleridae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 12: 45-49. 

Lis, J. A. 1996. Coleocydnus madagascariensis n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 7(3): 347-350. 

Lis, J. A. 1996. Some taxonomic changes in Afrotropical Cydnidae and new records for eight species of the family (Hemiptera, Heteroptera). Revue française d'Entomologie (ns) 18(3): 99-102. 

Lis, J. A. 1997. Taxonomy and phylogeny of the genus Cydnus Fabricius, 1803 (Hemiptera: Heteroptera: Cydnidae). Annales de la Sociéte entomologique de France 32(4)[1996]: 403-409. 

Lis, J. A. 1997. Macroscytus howei sp. n. (Heteroptera: Cydnidae) from the Lord Howe Island in the Tasman Sea. Polskie Pismo Entomologiczne 66(1-2): 27-32. 

Lis, J. A. 1997. Plataspidae, Thyreocoridae i Cydnidae. Keys for the identification of Polish insects, Part XVIII . True-bugs – Heteroptera 12: 28 pp. [in Polish] 

Lis, J. A. and A. Kocorek. 1997. The hindwing venation and its taxonomic value in Dinidoridae (Hemiptera: Heteroptera). International Journal of Invertebrate Taxonomy - Genus 8(3-4): 567-581. 

Lis, J. A. 1997. Studies on Cydnidae of the Australian Region. X. Two new species of the genus Chilocoris Mayr from New Guinea (Hemiptera: Heteroptera). International Journal of Invertebrate Taxonomy - Genus 8(3-4): 583-589. 

Lis, J. A. 1997. Studies on Cydnidae of the Australian Region. XI. A review of species from New Caledonia, with descriptions of Chilocoris neocaledonicus n. sp. and Macroscytus matilei n. sp. (Hemiptera: Heteroptera). International Journal of Invertebrate Taxonomy - Genus  (3-4): 591-602. 

Lis, J. A. 1997. Three new Australian genera of burrower bugs with four new species (Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 66: 189-201. 

Lis, J. A. 1997. Afroscytus, a new burrower bug genus from South Africa (Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 66: 203-209. 

Lis, J. A. 1997. Some remarks on the Palaearctic Cydnidae (Hemiptera: Heteroptera). Acta Entomologica Silesiana 4(1-2)[1996]: 3-7. 

Lis, J. A. 1998. An annotated checklist of burrower bugs (Cydnidae) of Thailand. Amemboa – News and Results on Heteroptera of Thailand 2: 43-46. 

Lis, J. A. 1998. Burrower bugs of the World (Heteroptera: Cydnidae) – the present state of knowledge. Proceedings of the Vth European Congress of Entomology 1: 307. České Budějovice, Czech Republic, August 23-29. 

Lis, J. A. 1998. Amnestus raunoi n. sp. from Iran, the first Old World representative of the New World subfamily Amnestinae (Hemiptera: Heteroptera: Cydnidae). Entomologica Fennica 9: 5-7. 

Lis, J. A. and B. Lis. 1998. True-bugs of Upper Silesia – an annotated checklist (Hemiptera: Heteroptera). Annals of the Upper Silesian Museum, Entomology 8-9: 107-146. 

Lis, J. A. and B. Lis. 1998. Acanthosomatidae, Scutelleridae. Keys for the identification of Polish insects, Part XVIII . True-bugs – Heteroptera 13: 32 pp. [in Polish] 

Lis, J. A. 1999. A revision of Australian species of the genus Macroscytus Fieber (Hemiptera: Heteroptera: Cydnidae). Entomologica scandinavica 29(4)[1998]: 459-479. 

Lis, J. A. 1999. Blaena tamasi sp. n. from Western Australia, the first species of Cydnidae (Hemiptera: Heteroptera) with staphylinoid wing modification. European Journal of Entomology 96: 53-55. 

Lis, J. A. 1999. Geopeltus neocaledonicus n. sp. from New Caledonia (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 10(1): 29-31. 

Lis, J. A. 1999. Macroscytus unisetosus n. sp., the first species of the javanus-group from New Guinea (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 10(1): 33-36. 

Lis, J. A. 1999. A revision of the Australian genus Eulonips Lis, with descriptions of three new species (Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 68(1): 29-39. 

Lis, J. A. 1999. Burrower bugs of the Old World – a catalogue (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy - Genus 10(2): 165-249. 

Lis, J. A. 1999. Taxonomy and phylogeny of Cephalocteinae with a reference to their historical biogeography (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 68(2): 111-131. 

Lis, J. A. 1999. The genus Chilocoris (Heteroptera: Cydnidae) in Australia. Acta Societatis Zoologicae Bohemicae 63: 139-155. 

Lis, J. A. 1999. Revision of the genus Macroscytus (Heteroptera: Cydnidae) in Madagascar and adjacent islands, with description of four new species. European Journal of Entomology 96: 427-437. 

Kocorek, A. and J. A. Lis. 2000. A cladistic revision of the Megymeninae of the World (Hemiptera: Heteroptera: Dinidoridae). Polskie Pismo Entomologiczne 69: 7-30. 

Lis, J. A., M. Becker, and C. W. Schaefer. 2000. Burrower Bugs (Cydnidae).  Pp. 405-419, in: Schaefer C. W. and A. R. Panizzi (eds), Heteroptera of Economic Importance. CRC Press, Boca Raton, London, New York, Washington D.C., 828 pp. 

Lis, J. A. 2000. A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy – Genus 11(3): 359-509. 

Lis, J. A. 2000. Cydnotomus - a new Australian burrower bug genus, with two new species (Heteroptera, Cydnidae). Deutsche Entomologische Zeitschrift 47: 137-141. 

Lis, J. A. 2000. Shield-bugs – Pentatomidae. Keys for the identification of Polish insects, Part XVIII . True-bugs – Heteroptera 14: 76 pp. [in Polish] 

Lis, J. A. 2000. Schiodtella africana sp. n. (Hemiptera: Heteroptera: Cydnidae), the first African representative of the genus. Polskie Pismo Entomologiczne 69(3): 263-266. 

Lis, J. A. 2000. A review of the Australian species of Geotomus Mulsant et Rey, 1866, with descriprions of three new species (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 69(4): 411-428. 

Lis, J. A. and A. Hohol. 2001. Geotomus garsaurioides, a peculiar new species of burrower bugs from Australia (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy – Genus 12(1): 7-12. 

Lis, J. A. 2001. Byrsinus setosus n. sp. from Flores and new records of some other Old World Cydnidae (Hemiptera: Heteroptera). International Journal of Invertebrate Taxonomy – Genus 12(1): 13-22. 

Lis, J. A. 2001. Prolactistes australis gen. et sp. nov. from Australia, with remarks on its systematic position within the family (Hemiptera: Heteroptera: Cydnidae). Annales Zoologici 51(1): 103-107. 

Lis, J. A. 2001. A review of the genus Byrsinus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae) in Australia, with a key to Australo-Pacific species. Annales Zoologici 51(2): 197-203. 

Lis, J. A. and A. Hohol-Kilinkiewicz. 2001. Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cephalocteinae (Hemiptera: Heteroptera: Cydnidae). Annales Zoologici 51(3): 293-297. 

Lis, J. A. 2001. Two new Oriental species of the burrower bug genus Chilocoris Mayr (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy – Genus 12(3): 269-274. 

Lis, J. A. 2001. European Cydnidae – the present state of knowledge (Heteroptera, Pentatomoidea). Program and Book of Abstracts. Lectures: 3. 2nd European Hemiptera Congress, Slovenia, Fiesa, 20-24 June, 2001.

Lis, J. A. and J. Heyna. 2001. Revision of the Australian genus Blaena (Heteroptera: Cydnidae). European Journal of Entomology 98(3): 321-350. 

Lis, J. A. 2001. Review of the species of Coleocydnus J. A. Lis, with a description of C. sicardi n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy – Genus 12(4): 429-436. 

Lis, J. A. and B. Lis. 2001. Cyclopelta robusta, a new species of dinidorid bugs (Hemiptera: Heteroptera: Dinidoridae) from Zaire. Polskie Pismo Entomologiczne 70(2): 147-150. 

Lis, J. A. and J. Heyna. 2001. Metathoracic wings venation in Cydnidae (Hemiptera: Heteroptera) and its bearing on the classification of the family. Annales Zoologici 51(4): 429-465. 

Lis, J. A. and A. Hohol-Kilinkiewicz. 2001. Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cydnidae (Hemiptera: Heteroptera). II. Amnestinae and Garsauriinae. Annales Zoologici 51(4): 467-471.

Lis, J. A. and J. Heyna. 2001. The metathoracic wing stridulitrum of the Cydnidae (Hemiptera: Heteroptera). Polskie Pismo Entomologiczne 70(4): 221-243. 

Lis, J. A. 2002. Megacydnus secundus sp. n. (Hemiptera: Heteroptera: Cydnidae) from the Democratic Republic of Congo. Polskie Pismo Entomologiczne 71(1): 3-6. 

Lis, J. A. 2002. Burrower bugs described after the Old World catalogue of the family (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 71(1): 7-17. 

Lis, J. A. 2002. The mesothoracic wing and its phylogenetic significance in Cydnidae (Hemiptera: Heteroptera: Pentatomoidea). Polskie Pismo Entomologiczne 71(1): 43-71. 

Lis, J. A. and D. Pluot-Sigwalt. 2002. Nymphal and adult cephalic chaetotaxy of the Cydnidae (Hemiptera: Heteroptera), and its adaptive, taxonomic and phylogenetic significance. European Journal of Entomology 99(1): 99-109. 

Lis, J. A. 2002. Peltoxys pilosus n. sp. from the Democratic Republic of Congo, with a key to known species of the genus (Hemiptera: Heteroptera: Cydnidae: Garsauriinae). International Journal of Invertebrate Taxonomy – Genus 13(2): 171-175. 

Lis, J. A. 2002. A redescription of Schiodtella laevicollis (Montandon) n. comb. (Hemiptera: Heteroptera: Cydnidae). International Journal of Invertebrate Taxonomy – Genus 13(2): 177-181. 

Lis, J. A. 2002. The family Cydnidae – a need for reclassification?. Second Quadrennial Meeting of the International Heteropterists’ Society, St. Petersburg 2002, Abstracts: 37. 

Lis, J. A. and A. Hohol-Kilinkiewicz. 2002. Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cydnidae (Hemiptera: Heteroptera). IV. Cydninae. Polskie Pismo Entomologiczne 71(2): 133-143. 

Lis, J. A., M. Jastrzębska, and A. Kocorek. 2002. Comparative studies on the pretarsal structures in Dinidoridae (Hemiptera: Heteroptera: Pentatomoidea). Polskie Pismo Entomologiczne 71(2): 165-184. 

Lis, J. A. and A. Hohol-Kilinkiewicz. 2002. Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cydnidae (Hemiptera: Heteroptera). III. Sehirinae. Annales Zoologici 52(3): 359-362. 

Lis, J. A. 2002. Notes on the South African genus Dearcla Signoret, 1883 (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 71(3): 265-271. 

Czaja, J. and J. A. Lis. 2002. Four new species of the Afrotropical genus Coleocydnus J. A. Lis (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 71(3): 273-292. 

Lis, J. A. and A. Hohol-Kilinkiewicz. 2002. Adult dorso-abdominal scent glands in the burrower bugs (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 71(4): 359-395. 

Lis, J. A. 2003. A redescription of Byrsinus nigroscutellatus (Montandon) comb. n. (Hemiptera: Heteroptera: Cydnidae), with remarks on the synonymy of Byrsinocoris Montandon, 1900 and Byrsinus Fieber, 1860. Polskie Pismo Entomologiczne 72(1): 39-4. 

Lis, J. A. 2003. Garsauriella hirta sp. n. (Hemiptera: Heteroptera: Cydnidae) from the Democratic Republic of Congo. Polskie Pismo Entomologiczne 72(1): 45-49. 

Lis, J. A. 2003. Dearcla congoensis, a new species of burrower bugs (Hemiptera: Heteroptera: Cydnidae) from the Democratic Republic of Congo. Polskie Pismo Entomologiczne 72(2): 139-143. 

Czaja, J. and J. A. Lis, J. A. 2003. Coleocydnus inexpectatus, a new Afrotropical species of burrower bugs (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 72(2): 197-202. 

Lis, J. A. 2003. Macroscytus affinis sp. n. from Mindanao Island in the Philippines (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 72(3): 215-218. 

Lis, J. A. 2003. External morphology of the tympanal organs of Dinidoridae (Hemiptera: Heteroptera: Pentatomoidea). Polskie Pismo Entomologiczne 72(4): 297-304. 

Lis, J. A. 2003. Ovaries and lateral oviducts of the female internal reproductive system in five specis of burrower bugs (Hemiptera: Heteroptera: Cydnidae). Polskie Pismo Entomologiczne 72(4): 305-312. 

Lis, J. A. and C. W. Schaefer. 2005. Tibial combs in the Cydnidae (Hemiptera: Heteroptera) and their functional, taxonomic and phylogenetic significance. Journal of Zoological Systematics and Evolutionary Research, 43 (4): in press.

Lis, J. A. 2006. Heissocteus ernsti nov. gen. et nov. sp. (Heteroptera, Cydnidae) from Zambia. In: Rabitsch, W. (ed.), Hug the Bug - For Love of True Bugs. Festschrift zum 70. Geburtstag von Ernst Heiss. Denisia 19: 505-507.

Lis, J. A. and B. Lis. 2007. Peltoxys sataranus, a new burrower bug species from India, and new records for some other heteropterans from the Maharashtra State (Hemiptera: Heteroptera). Genus 18(2): 209-214.

Lis, J. A. and M. Webb. 2007. Redescription of the burrower bug Adrisa sepulchralis (Erichson, 1842) (Insecta: Hemiptera: Cydnidae), based on the only known male (recently introduced to the UK from Australia), and the lectotype from Tasmania. Entomologist’s Monthly Magazine 143: 59-65.

Kocorek, A. and J. A. Lis. 2008. Megymenum sumatranum, a new dinidorid species from Sumatra (Hemiptera: Heteroptera: Dinidoridae. Genus 19(1): 21-25.

Lis, J. A. and B. Lis. 2008. Lactistes obesipes Signoret, 1879 (Hemiptera: Cydnidae) – the burrower bug enigma: Redescription, synonym, and remarks on its distribution and taxonomic status. Zootaxa 1933: 66-68.

Pluot-Sigwalt, D. and J. A. Lis. 2008. Morphology of the spermatheca in the Cydnidae (Hemiptera: Heteroptera): Bearing of its diversity on classification and phylogeny. European Journal of Entomology 105(2): 279-312.

Lis, J. A. 2010. Coxal combs in the Cydnidae sensu lato and three other related "cydnoid" families – Parastrachiidae, Thaumastellidae, Thyreocoridae (Hemiptera: Heteroptera): Functional, taxonomic, and phylogenetic significance. Zootaxa 2476: 53-64.

Lis, J. A. 2010. Pretarsal structures in the family Parastrachiidae (Hemiptera: Heteroptera: Pentatomoidea). Zootaxa 2693: 60-62.

Lis, J. A. and B. Lis. 2010. Teabooma secunda (Hemiptera: Heteroptera: Cydnidae) – a new species of a New Caledonian endemic genus, with a redescription of T. princeps Distant. Zootaxa 2543: 64-68.

Lis, J. A. and R. S. Zack. 2010. A review of burrower bugs (Hemiptera: Heteroptera: Cydnidae sensu lato) of Guam. Zootaxa 2523: 57-64.

Lis, J. A. and D. J. Ziaja. 2010. Pretarsal structures in the family Cydnidae sensu lato (Hemiptera: Heteroptera: Pentatomoidea). Zootaxa 2545: 23-32.

Lis, J. A. 2011. Description of the unknown male of the Afrotropical Heissocteus ernsti J. A. Lis (Hemiptera: Heteroptera: Cydnidae: Cephalocteinae). Zootaxa 2762: 61-62.

Lis, J. A. and B. Lis. 2011. Is accurate taxon identification important for molecular studies? Several cases of faux pas in pentatomoid bugs (Hemiptera: Heteroptera: Pentatomoidea). Zootaxa 2932: 47-50.

Kędzior, M., J. A. Lis, and A. Wolski. 2012. Terrestrial true-bugs (Hemiptera: Heteroptera) of the Czantoria Wielka massif (Silesian Beskid Mts.). Heteroptera Poloniae – Acta Faunistica 5: 17-24.

Lis, J. A., P. Lis, D. J. Ziaja, and A. Kocorek. 2012. Systematic position of Dinidoridae within the superfamily Pentatomoidea (Hemiptera: Heteroptera) revealed by the Bayesian phylogenetic analysis of the mitochondrial 12S and 16S rDNA sequences. Zootaxa 3423: 61-68.

Lis, J. A., J. Olchowik, and M. Bulińska-Balas. 2012. Preliminary studies on the usefulness of DNA mini-barcodes for determining phylogenetic relationships within shieldbugs (Hemiptera: Heteroptera: Pentatomidae). Heteroptera Poloniae – Acta Faunistica 4: 13-25.

Kłys, G. and J. A. Lis. 2013. Chilocoris nitidus Mayr, 1865, the first Oriental burrower bug recorded in Tajikistan, with remarks on the zoogeography of the nitidus-group of Chilocoris Mayr, and an annotated checklist of Tajik Cydnidae (Hemiptera: Heteroptera). Zootaxa 3641(2): 193-196.

Kłys, G. and J. A. Lis. 2013. First cave records for Palearctic burrower bugs (Hemiptera: Heteroptera: Cydnidae) from Tajikistan, with a checklist of the World Cydnidae associated with caves. Zootaxa 3686(4): 493-496.

Lis, J. A. 2013. Family Cydnidae Billberg, 1820. Pp. 132-135, in: Gerlach, J. (ed.), Odonata, Hemiptera, Hymenoptera and Other Insects of the Seychelles Islands. Siri Scientific Press, Manchester, UK, 394 pp.

Lis, J. A. and B. Lis. 2014. First record of Megacydnus secundus J. A. Lis, 2002, a representative of Afrotropical endemic burrower bug genus from Uganda, and an annotated checklist of Ugandan Cydnidae (Hemiptera: Heteroptera). Zootaxa 3795(4): 494-496.

Lis, J. A., B. Lis, D. J. Ziaja, and R. Dobosz. 2014. Description and DNA barcoding of Ochetostethomorpha secunda, a new species of the South African endemic burrower bug genus (Hemiptera: Heteroptera: Cydnidae) from Namibia. Zootaxa 3884(6): 561-566.

Lis, J. A., B. Lis, D. J. Ziaja, and Y. Nakatani. 2014. Towards resolving a problem of the identity of the Aethus species (Hemiptera: Heteroptera: Cydnidae) occurring in Cambodia. Zootaxa 3895(3): 446-450.

Lis, J. A. and D. J. Ziaja. 2014. New records of Namibian burrower bugs (Heteroptera: Pentatomoidea: Cydnidae). Heteroptera Poloniae - Acta Faunistica 8(1): 5-8.

Lis, J. A. 2015. Chilocoris quadraticollis Linnavuori, 1993 (Hemiptera: Heteroptera: Cydnidae): first records from the Democratic Republic of Congo with first data on its biology. Heteroptera Poloniae – Acta Faunistica 9: 43-44.

Lis, J. A., A. Kocorek, D. J. Ziaja, and P. Lis. 2015. New insight into the systematic position of the endemic Madagascan genus Amberiana (Hemiptera: Heteroptera: Dinidoridae) using 12S rDNA sequences. Turkish Journal of Zoology 39: 610-619.

Lis, J. A. and B. Lis. 2015. Raunoloma longiceps (Linnavuori, 1977) (Hemiptera: Heteroptera: Cydnidae): first record from Uganda. African Entomology 23(1): 255-256.

Lis, J. A. and D. J. Ziaja. 2015. An annotated checklist of burrower bugs (Hemiptera: Heteroptera: Cydnidae) from Bali (Indonesia) with new records. Polskie Pismo Entomologiczne 84: 305-310.

Vršanský, P., J. A. Lis, J. Schlögl, M. Guldan, T. Mlynský, P. Barna, and P. Štys. 2015. Partially disarticulated new Miocene burrower bug (Hemiptera: Heteroptera: Cydnidae) from Cerová (Slovakia) documents occasional preservation of terrestrial arthropods in deep-marine sediments. European Journal of Entomology 112(4): ???.

Lis, B. and J. Lis. 2016. Chilocoris laevicollis Horváth, 1919, and Ch. umbricola Linnavuori, 1993 – two trogloxenic burrower bugs recorded for the first time in Gabon (Central Africa). Zootaxa 4061(3): 286-290.

Lis, J. A. 2016. Nishadana typica (Distant, 1889) (Hemiptera: Heteroptera: Cydnidae): first record from Laos. Heteroptera Poloniae – Acta Faunistica 10: 55-56.

Lis, J. A. and B. Lis. 2016. Macroscytus raunoi, a new burrower bug species from Papua New Guinea (Hemiptera: Heteroptera: Cydnidae: Cydninae). Entomologica Americana 122(1-2): 38-41.

Lis, J. A. and B. Lis. 2016. Chilocoris serratus n. sp., the first burrower bug species (Hemiptera: Heteroptera: Cydnidae) recorded in Guinea-Bissau with an annotated checklist of the Afrotropical species of the genus Chilocoris Mayr, 1865. Zootaxa 4103(4): 383-389.

Lis, J. A. and B. Lis. 2016. Amnestus mendeli sp. n., the first burrower bug species (Hemiptera: Heteroptera: Cydnidae) recorded on Ascension Island, central Atlantic Ocean. Zootaxa 4126(1): 115-122.

Lis, J. A., B. Lis, and S. G. Compton. 2016. Chilocoris capensis n. sp., the first species of the genus Chilocoris Mayr, 1865 (Hemiptera: Heteroptera: Cydnidae) recorded in the Republic of South Africa with an annotated checklist of South African burrower bugs. Zootaxa 4147(5): 564-574.

Lis, J. A., B. Lis, and D. J. Ziaja. 2016. In BOLD we trust? A commentary on the reliability of specimen identification for DNA barcoding: a case study on burrower bugs (Hemiptera: Heteroptera: Cydnidae). Zootaxa 4114(1): 83-86.

Coscarón, M. del C. and J. Lis. 2017. Family Cydnidae Billberg. Pp. 83-88, in: Coscarón, M. del C. (ed.), A catalogue of the Heteroptera (Hemiptera) or true bugs of Argentina. Zootaxa 4295: 1-432.

Lis, J. A., D. J. Ziaja, B. Lis, and P. Gradowska. 2017. Non-monophyly of the “cydnoid” complex within Pentatomoidea (Hemiptera: Heteroptera) revealed by Bayesian phylogenetic analysis of nuclear rDNA sequences. Arthropod Systematics & Phylogeny 75(3): 481-496.

Lis, J. A., B. Lis, and E. Heiss. 2018. Chilamnestocoris mixtus gen. et spec. nov., the first burrower bug (Hemiptera: Pentatomoidea: Cydnidae) in Upper Cretaceous Burmese amber. Cretaceous Research 91: 257-262.

Lis, J. A. and P. F. Whitehead. 2019. Another alien bug in Europe: the first case of transcontinental introduction of the Asiatic burrower bug Macroscytus subaeneus (Dallas, 1851) (Hemiptera: Heteroptera: Cydnidae) to the U.K. through maritime transport. Zootaxa 4555(4): 588-594.

Lis, J. A., M. Roca-Cusachs, B. Lis, and S. Jung. 2020. Pullneyocoris dentatus gen. et sp. nov. (Hemiptera: Pentatomoidea: Cydnidae), the third representative of the subfamily Amnestinae from mid-Cretaceous amber of northern Myanmar. Cretaceous Research 115(104562): 1-8.

Lis, J. A. and A. Zielińska. 2020. Review of the genus Prolactistes (Heteroptera: Cydnidae), with two new combinations, first Indonesian records and a key to species of the genus. Acta Entomologica Musei Nationalis Pragae 60(2): 609-614.

Lis, J. A., and B. Lis. 2021. Geotomus granulosus, a peculiar sehirine-like new species of the subfamily Cydninae (Hemiptera: Heteroptera: Cydnidae) from Burundi. Zootaxa 5087(3): 489-493.

Lis, J. A. 2023. Systematic position of the fossil burrower bug Eocenocydnus lisi Popov, 2019 (Hemiptera, Heteroptera, Cydnidae) revealed by a parallel/cross-eyed viewing method used for obtaining three-dimensional images. Insects 14(22): 8 pp.

Lis, J. A. 2023. Molecular apomorphies in the secondary and tertiary structures of length-variable regions (LVRs) of 18S rRNA shed light on the systematic position of the family Thaumastellidae (Hemiptera: Heteroptera: Pentatomoidea). International Journal of Molecular Sciences 24(7758): 22 pp.

Lis, J. A. and P. J. Domagała. 2024. Inconsistencies in the classification of the family Cydnidae (Hemiptera: Heteroptera: Pentatomoidea) revealed by molecular apomorphies in the secondary and tertiary structures of 18S rRNA length-variable region L (LVR L). International Journal of Molecular Sciences 25(939): 27 pp.

 

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David A. Rider
Professor of Entomology
North Dakota State University
E-Mail: David.Rider@ndsu.edu

updated: 17 January 2024 

Published by the Department of Entomology 


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